The Howler is most effective when supporting allies from a distance. Its indirect fire allows it to attack enemies on cliffs and mountains.
The Howler is the only tier-1 unit to deal area-of-effect damage.
Use ground targeting to aim the Howler's attack toward where you anticipate enemies are going—not where they are at the time.
Howlers excel at countering smaller tier-1 units, but lose potency against larger enemies.
The Howler's range and area-of-effect radius increase, but its rate of fire decreases.